These creatures are commonly present in a pollen slide from the Early Pleistocene in Holland, which flora is dominated by Picea and Pinus with some Tsuga.
see Brown & Hebda 2011 Origin, development, and dynamics of coastal temperate conifer rainforests of southern Vancouver Island, Canada.Canadian Journal of Forest Research 32(2):353-372.
These could be foraminiferal organic linings (see attached paper by Tyszka et al, 2021), which have been reported in several instances from Quaternary palynological preparations by Bas van Geel and others.
Indeed, but is the irregularity real, or only apparent (due to taphonomy, sample processing, etc.). I don't know what is their size, but they may be too big to be typical fungal material. Their sizes and how they compare to those of foraminiferal linings may solve this. Especially the one at the top right in your montage reminded me of the coiled foraminiferal linings, then the others could be explained as disrupted such specimens. I would also try to contact Bas van Geel, who has worked extensively on non-pollen palynomorphs, and Tiszka, whose paper I had attached previously). I am attaching another paper that may come in handy.
Bas van Geel supports your interpretation that these could represent foram linings. Thanks again, Alexandru. I will resample the surface section. This is worth more research.