The reason is rapid activation of the avian ventricular myocardium via abundant Purkinje system which delivers impulse not only to the subendocardium (as in the majority of mammals) but to the intramural layers or even subepicardium. This specific activation pattern results in the specific ECG features. For details, you could refer to the paper of my colleague: Kharin SN. Depolarisation and repolarisation sequences of ventricular epicardium in chickens (Gallus gallus domesticus). Comp Biochem Physiol A Mol Integr Physiol. 2004 Jan;137(1):237-44. PubMed PMID: 14720609. Also you may search through PubMed for the papers by Shmakov DN and coauthors concerning myocardial activation in different verterbrates.