I am working on multi species occupancy modelling and I stumbled upon creating residual covariance matrix which has a range of -1 to +1. I need to get a very basic understanding of what a residual covariance matrix is showing me? Is it as simple as showing relation between each species? So, if the value comes to be negative between two species, does it mean they have a negative relation? I am confused because of the term "residual covariance". I would appreciate guidance on this.

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