Low temperatures could completely damage the flowers or injure developing fruit tissue resulting in misshapen fruit. Pollination is also affected if temperature drops below 60 oF (15 oC) during flowering. As a result, some achenes do not have a seed, remain small, and cause fruit distortion. Cold injury can also cause fruit with multiple tips. Distorted fruit from lygus bug damage have uniform sized achenes, but the achenes affected by cold injury are much smaller than those on unaffected parts of the fruit. (cited in the above link)
Proteomic study of low-temperature responses in strawberry cultivars (Fragaria x ananassa) that differ in cold tolerance. Plant physiology 06/2012; 159(4):1787-805.
Article Proteomic Study of Low-Temperature Responses in Strawberry C...
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