Let’s derive a general formula for the equilibrium allele frequencies in the case where the heterozygote is either more fit or less fit than either homozygote.  For the case of heterozygote superiority, the selection coefficients, r and s, are negative numbers (‐1   0). Show that the frequency of a in the next generation of adults is q' = (q(1+sq))/(1+r(1-q)^2+sq^2).   Show that the steady‐state solutions of this discrete dynamical system are: q = 0, q = 1, and q = r/(s+r).  

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