some ideas were published at https://www.researchgate.net/post/Why_are_plant_meristems_always_virus_free
Certainly, this is not the original citations:
1) In the meristem there is no cells differentiation. Virus can't move from cell to cell through plasmodesmesta using their Movement protein.( Ghislain Kanfany)
2) the phloem is not enough developed in the meristem, so that the virus can't spread very quickly at the meristem. ( Mathieu Pierre)
3) there is a very intense metabolic activity in the meristem (Mathieu Pierre)
4) viral replication (viral inactivating system) cannot cope with faster meristem cell division and auxin level might also limit viruses ( Kamran Iqbal Shinwari)
5) An RNA-dependent RNA polymerase prevents meristem invasion by viruses (Schwach F, Vaistij FE, Jones L, Baulcombe DC. An RNA-dependent RNA polymerase
prevents meristem invasion by potato virus X and is required for the activity but not the production of a systemic silencing signal. Plant Physiol. 2005 Aug;138(4):1842-52.
some more mechanisms were discussed...
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