What are the evolutionary origins of the vitamin B12 molecule?

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Scott Ferrenberg

Not sure if this article is answering your question, but it describes the independent evolution, and likely selective driver of B12 synthesis in microbial taxa.

http://mbe.oxfordjournals.org/content/28/10/2921.full

Derek Townshend

Thanks Scott. That is exactly the type of background information I am looking for.