I was trying to simulate this system from papers attached here. But it is not working. I am attaching my code. Please see it.

As a first step I am trying to simulate the spiral wave patterns generated by extracellular cAMP secreted by Dd. The system has the set of equations which has 4 ODEs and 1PDE. And I am also doubtful regarding the implementation of boundary conditions in this program (Neumann no flux boundary condition). I am attaching the equation list too.

http://www-users.math.umn.edu/~othmer/papers/Tang:1995:EOW.pdf

http://www-users.math.umn.edu/~othmer/papers/Dallon:1997:DCM.pdf

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