I try to search litrerature about this subject but there are a few publications about it. But I find this artickle(attach file). I hope it will be helpful. Best regards
Shiju T. Raj Thomas K. Sabu Zhao Danyang (2012) The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent : taxonomy with notes on habits and distributional patterns
Page 1367 in: Yves Bousquet. 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico PenSoft Publishers LTD, Pp.1722. ISBN954642658X, 9789546426581
The apterous endemic genus Omphra Dejean (Coleoptera: Carabidae: Helluonini) of the Indian subcontinent: taxonomy with notes on habits and distributional patterns, Shiju T. Raj, Thomas K. Sabu, and Zhao Danyang