Can you help me with this simbiology error- Invalid assignment syntax or parse error for rule- ?

12 December 2013 0 7K Report

I am exercising on the article of Tzafriri et al(2004) titled "Analysis of compartmental models of ligand induced endocytosis" with simbiology. But I am taking the error above according to my rate rules. I think my expressions can be wrong, but I don't know how to correct them. Can you please help me?

My simbiology project file is attached below.

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