I am not an expert of African languages, but I know that in INALCO-Paris someone study those languages in a comparative perspective https://scholar.google.com/citations?view_op=view_citation&hl=fr&user=_FPXWaoAAAAJ&citation_for_view=_FPXWaoAAAAJ:IWHjjKOFINEC
even this website could be useful to look for interesting references https://wals.info/languoid/lect/wals_code_hau