The direct flight musculature of dragonflies comprises of stroke frequency, amplitude, phase and angle of attack can be varied independently on each of the four wings. Dragonflies put these abilities to excellent effect, using them to enlist a variety of wing kinematics in free flight.
Alongwith these, The direct flight musculature and four-winged morphology of dragonflies make them ideal candidates for exploiting such aerodynamic effects.
Elongated two pairs of wings of a dragonfly are coordinated with strong muscle for helping them in their forward as well as backward movements. While flying backwards, the wings are so twisted that they start pushing air in the forward direction and help them to flew backward.