I found Copris lunaris female on a meadow near a cow dung, 2-3 days old. It is near a meadow marshes, where people come frequently with animals for food. I found the specimen in May and the temperature was 21°C.
This won't help much for the determination but if (and it's a big if !) this is a Mesostigmata, I recommend this interesting paper about species associated with Copris lunaris :
http://www.eje.cz/pdfs/eje/2009/04/11.pdf
Good luck, many mites are difficult to identify unless you have much experience.
The second and third photos are (may be are similar) females of Macrocheles genus (Macrochelidae Family), may be M. muscaedomestacae species! but I am not sure about the species, because the photoes are unclare.
I'm not happy with the quality of the photos. Next week I will go to a laboratory with stereo-microscope more powerful and hope to be more clear, to be observe some criteria for determination. I know the photo is difficult to determine, but try .... Can I send these photos, the following week?
Ok, please take some photographs on sternal shield ornamentation (with high magnification). if possible, I would like to know which dorsal setae are pilose!
I agree : Macrocheles sp. and Uropodid. On Copris several species can be found In European countries most of the species can be determined using the key from "A review of the Macrochelidae (Acari: Mesostigmata) of the British Isles", (download at "http://www.biodiversitylibrary.org/part/17598#/summary" with illustrations, or "Review of the phoretic association between coprophilous insects and macrochelid mites (Acari: Mesostigmata) in France" Niogret et al. Elytron (key). Available on Researchgate. However the mite must be re-hydrated (you can use heated lactic acid to help prompt recovery!)
Many heterostigmatic mites including speciose superfamily Pygmephoroidea and also parasitic mites of the family Podapolipidae may be present on this beetle and other close and sympatric beetles.