Birkhead in his book "Bird Sense" (2012) writes that unfortunately there is very little published about touch receptors in the bills of small birds.

He also refers to the work of Herman Berkhoudt (1980, 1985)  who has studied this issue and who found several touch receptors including Merkel cell receptors, double-column Merkel cell receptors and many Herbst corpuscles, all those in a zebra finch bill (Birkhead 2012). Other species beaks may contain even more receptors. I also tend to believe that both cormorants and pelicans may well have lots of such yet unstudied receptors in their beaks or pouches or other parts of their front bodies which help them locate prey. Unfortunately, I suppose there is no or very little research done in such anatomical and physiological issues on birds right now, so we may discover them after many years.

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