I have recently been looking up data on the extinct giant (hummingbird-sized) Eocene ant Titanomyrma, as well as the closely related and possibly synonymous form genus Formicium. However, the only information I have been able to find on the paleobiology of these species is from Wikipedia, and none of the information they give cites a particular paper.

According to Wikipedia, Titanomyrma lacks a closing mechanism on the crop (whatever that means), sprayed formic acid as a primary means of defense, and has adaptations that suggest it was either a fungivore like modern leaf-cutter ants or was predatory in a manner similar to driver ants. Does anyone know what research articles (if any) proposed these ideas, and why? It could be that they were proposed in a paper regarding Formicium (which species assigned to Titanomyrma were formerly placed in).

Finally, as an additional question, does anyone know what living group of ants the Formiciinae (note the extra "i") are related to? The group may be extinct, but close living relatives are known it might be easier to determine how likely the presence/absence of these adaptations are.

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