Carbohydrate polymerization is a flexible tool mainly for osmoregulation of water intake. The main pressure control is exrted by vacuole sap composition.
A ref on H tuberosus is added https://www.researchgate.net/publication/233803968_Localization_of_Fructan_Metabolism_in_the_Vacuoles_Isolated_from_Protoplasts_of_Jerusalem_Artichoke_Tubers_Helianthus_tuberosus_L
Article Localization of Fructan Metabolism in the Vacuoles Isolated ...